TYPE: Costa Rica. Border of San José and Cartago provinces: Pan-American Highway, km 52. El Empalme, c. 300 m after the turning to Santa María de Dota, 09º42'49"N, 83º56'58"W, 26 Jul 2003, W.M. Whitten 2164, M.A. Blanco & D. Bogarín (holotype, USJ; isotype, JBL-spirit; illustration of type).